|Photo by Ohio State Weed Lab, The Ohio State University, Bugwood.org|
|Division:||Magnoliophyta - Flowering plants|
|Class:||Magnoliopsida – Dicotyledons|
| Physalis heterophylla|
|Natural range of Physalis heterophylla from USDA NRCS Plants Database.|
Common name: Clammy groundcherry
Synonyms: Physalis heterophylla var. heterophylla; P. heterophylla var. ambigua (A. Gray) Rydberg; P. heterophylla var. clavipes Fernald; P. heterophylla var. nyctaginea (Dunal) Rydberg; P. ambigua (A. Gray) Britton; P. nyctaginea Dunal
"Annual or perennial, glabrous or pubescent herbs. Leaves petiolate, entire to coarsely dentate, the base tapered, subcordate or oblique. Flower axillary, solitary; the calyx tube expanding, papery and bluntly conical when mature, loosely enclosing the fruit. Corolla campanulate to subrotate, yellow, often with 5 dark spots in the base of the tube; stamens 5, erect, separate, anther yellow or blue, dehiscing by lateral slits berry somewhat mealy, 2-locular, globose." 
"Rhizomatous, often viscid perennial 2-9 dm tall. Stem pubescence of glandular trichomes mixed with both long and short nonglandular trichomes. Leaves widely ovate or oblanceolate to triangular or lanceolate, 3-11 cm long, 3-8 cm wide, sinuate-dentate or coarsely and irregularly dentate, rarely subentire, weakly cordate. Calyx 7-12 mm long, lobes triangular to lanceolate, shorter than the tube; corolla 10-18 mm long or broad, with 5 brownish basal spots; anthers yellow or rarely bluish, 3.5-4.5 mm long. Berry ca. 10 mm in diam., the fruiting calyx 2.5-3 cm long, about as broad." 
P. heterophylla has been documented to grow in calcareous coastal hardwood hammocks, drying sand bordering woods, upland sandhills and annually burned longleaf pineland.  It has also been observed growing in old biocontrol plots and a fallow quail food patch. It has been documented in moist sandy loam and in the loam soils of the Red Hills Region.  Associated species include longleaf pine. 
This species is thought to be dispersed by consumption by vertebrates. 
Conservation and management
Cultivation and restoration
References and notes
- Radford, Albert E., Harry E. Ahles, and C. Ritchie Bell. Manual of the Vascular Flora of the Carolinas. 1964, 1968. The University of North Carolina Press. 927. Print.
- Florida State University Robert K. Godfrey Herbarium database. URL: http://herbarium.bio.fsu.edu. Last accessed: July 2015. Collectors: Loran C. Anderson, Andre F. Clewell, R. F. Doren, R. Komarek, J. M. Kane, Lisa Keppner. States and Counties:Florida: Calhoun, Leon, Nassau, Wakulla, Washington. Georgia: Thomas. Compiled by Tall Timbers Research Station and Land Conservancy.
- Nelson, G. PanFlora: Plant data for the eastern United States with emphasis on the Southeastern Coastal Plains, Florida, and the Florida Panhandle. www.gilnelson.com/PanFlora/ Accessed: 12 DEC 2016
- Kirkman, L. Katherine. Unpublished database of seed dispersal mode of plants found in Coastal Plain longleaf pine-grasslands of the Jones Ecological Research Center, Georgia.