Difference between revisions of "Trichostema dichotomum"

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==Description==  
 
==Description==  
 
<!-- Basic life history facts such as annual/perrenial, monoecious/dioecious, root morphology, seed type, etc. -->
 
<!-- Basic life history facts such as annual/perrenial, monoecious/dioecious, root morphology, seed type, etc. -->
"Pubescent annual or weak perennial herbs from tap roots, freely branched with opposite branches. Stems obscurely angled, usually stipitate-glandular, leafy. Leaves entire or toothed, rarely lobed, petiolate to essentially sessile. Inflorescence a panicle of paired, bracteate, helicoid cymes with a flower at the axis of each pair; bracts similar to the leaves, smaller. Calyx bilabiate, lower lip longer than the upper with 3 essentially equal teeth, upper with 2 similar teeth; corolla blue to violet, zygomorphic, 5-lobed, lowest lobe lip-like, 5-10 mm long, lateral triangular-ovate, 2-4 mm long, tube short. Stamens 4, ca. 2X as long as the corolla, curved between the lateral lobes and curled downward almost meeting the lip; stigma bilobed, style curled in same fashion as stamens. Mericarps brownish to olive or blackish, dull, obovoid." - Radford et al 1964
+
"Pubescent annual or weak perennial herbs from tap roots, freely branched with opposite branches. Stems obscurely angled, usually stipitate-glandular, leafy. Leaves entire or toothed, rarely lobed, petiolate to essentially sessile. Inflorescence a panicle of paired, bracteate, helicoid cymes with a flower at the axis of each pair; bracts similar to the leaves, smaller. Calyx bilabiate, lower lip longer than the upper with 3 essentially equal teeth, upper with 2 similar teeth; corolla blue to violet, zygomorphic, 5-lobed, lowest lobe lip-like, 5-10 mm long, lateral triangular-ovate, 2-4 mm long, tube short. Stamens 4, ca. 2X as long as the corolla, curved between the lateral lobes and curled downward almost meeting the lip; stigma bilobed, style curled in same fashion as stamens. Mericarps brownish to olive or blackish, dull, obovoid." < ref name="Radford et al 1964">Radford, Albert E., Harry E. Ahles, and C. Ritchie Bell. Manual of the Vascular Flora of the Carolinas. 1964, 1968. The University of North Carolina Press. 897. Print.</ref>
  
"At least the principal stems spreading pubescent, the trichomes 0.2-1 mm long, usually bearded at the nodes. Leaves elliptic to elliptic-lanceolate, 2-7 cm long, 0.4-2.3 cm wide, entire, crenate, or rarely loved, base tapered to cuneate or attenuate; petioles 0-1.5 cm long. Mericarps distinctly reticulate, 1.9-2.3 mm long." - Radford et al 1964
+
"At least the principal stems spreading pubescent, the trichomes 0.2-1 mm long, usually bearded at the nodes. Leaves elliptic to elliptic-lanceolate, 2-7 cm long, 0.4-2.3 cm wide, entire, crenate, or rarely loved, base tapered to cuneate or attenuate; petioles 0-1.5 cm long. Mericarps distinctly reticulate, 1.9-2.3 mm long." <ref name="Radford et al 1964"/>
  
 
==Distribution==
 
==Distribution==
 
==Ecology==
 
==Ecology==
 
===Habitat=== <!--Natural communities, human disturbed habitats, topography, hydrology, soils, light, fire regime requirements for removal of competition, etc.-->
 
===Habitat=== <!--Natural communities, human disturbed habitats, topography, hydrology, soils, light, fire regime requirements for removal of competition, etc.-->
In the Coastal Plain in Florida and Georgia, ''T. dichotomum'' has been found in burned upland pinewoods; longleaf pine forests; annually burned pine savannas; loamy sand of woodlands; loamy sand of open pine-hickory woods; floodplain forests; sandy loam along bluffs; amidst cypress knees on the shores of lakes and ponds; sandy ridges; burned pond pine shrub bog flatwoods; longleaf pine-turkey oak sand ridges; and at the edges of an open limestone glade (FSU Herbarium). It has been recorded in disturbed habitats such as roadsides; deciduous flatwoods clearings; and a mowed meadow. Associated species include ''Vaccinium stamineum, Trichostema setaceum, Scoparia, Panicum, Andropogon gerardii, Angelica dentata'', and ''Helianthus radula'' (FSU Herbarium).
+
In the Coastal Plain in Florida and Georgia, ''T. dichotomum'' has been found in burned upland pinewoods; longleaf pine forests; annually burned pine savannas; loamy sand of woodlands; loamy sand of open pine-hickory woods; floodplain forests; sandy loam along bluffs; amidst cypress knees on the shores of lakes and ponds; sandy ridges; burned pond pine shrub bog flatwoods; longleaf pine-turkey oak sand ridges; and at the edges of an open limestone glade. <ref name="FSU Herbarium">Florida State University Robert K. Godfrey Herbarium database. URL: [http://herbarium.bio.fsu.edu http://herbarium.bio.fsu.edu]. Last accessed: July 2015. Collectors: Loran C. Anderson, Robert K. Godfrey, R. Komarek, R. A. Norris, Andre F. Clewell, K. Craddock Burks, David Dixon, O. Lakela, R. Kral, James P. Gillespie, R. L. Lazor, A. H. Curtiss, Roy Jervis, John B. Nelson, R. E. Perdue, Jr., Richard S. Mitchell, Bian Tan, Nancy E. Jordan, Ann F. Johnson, Wilson Baker, Cecil R Slaughter, Marc Minno. States and Counties: Florida: Baker, Calhoun, Citrus, Columbia, Duval, Franklin, Gadsden, Gulf, Jackson, Jefferson, Lafayette, Leon, Liberty, Madison, Okaloosa, Wakulla, Walton. Georgia: Grady, Thomas. Compiled by Tall Timbers Research Station and Land Conservancy.</ref> It has been recorded in disturbed habitats such as roadsides; deciduous flatwoods clearings; and a mowed meadow. Associated species include ''Vaccinium stamineum, Trichostema setaceum, Scoparia, Panicum, Andropogon gerardii, Angelica dentata'', and ''Helianthus radula.'' <ref name="FSU Herbarium"/>
  
 
===Phenology=== <!--Timing off flowering, fruiting, seed dispersal, and environmental triggers.  Cite PanFlora website if appropriate: http://www.gilnelson.com/PanFlora/ -->
 
===Phenology=== <!--Timing off flowering, fruiting, seed dispersal, and environmental triggers.  Cite PanFlora website if appropriate: http://www.gilnelson.com/PanFlora/ -->
It has been recorded flowering September and October, and fruiting September through December (FSU Herbarium).
+
It has been recorded flowering September and October, and fruiting September through December. <ref name="FSU Herbarium"/>
  
 
===Seed dispersal===
 
===Seed dispersal===
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<!--===Fire ecology===--> <!--Fire tolerance, fire dependence, adaptive fire responses-->
 
<!--===Fire ecology===--> <!--Fire tolerance, fire dependence, adaptive fire responses-->
 
===Pollination===
 
===Pollination===
The following Hymenoptera families and species were observed visiting flowers of ''Trichostema dichotomum'' at Archbold Biological Station (Deyrup 2015):
+
The following Hymenoptera families and species were observed visiting flowers of ''Trichostema dichotomum'' at Archbold Biological Station: <ref name="Deyrup 2015">Deyrup, M.A. and N.D. 2015. Database of observations of Hymenoptera visitations to flowers of plants on Archbold Biological Station, Florida, USA.</ref>
  
 
Halictidae:  ''Lasioglossum placidensis''
 
Halictidae:  ''Lasioglossum placidensis''
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==References and notes==
 
==References and notes==
Deyrup, M.A. and N.D. 2015. Database of observations of Hymenoptera visitations to flowers of plants on Archbold Biological Station, Florida, USA.
 
 
Florida State University Robert K. Godfrey Herbarium database. URL: [http://herbarium.bio.fsu.edu http://herbarium.bio.fsu.edu]. Last accessed: July 2015. Collectors: Loran C. Anderson, Robert K. Godfrey, R. Komarek, R. A. Norris, Andre F. Clewell, K. Craddock Burks, David Dixon, O. Lakela, R. Kral, James P. Gillespie, R. L. Lazor, A. H. Curtiss, Roy Jervis, John B. Nelson, R. E. Perdue, Jr., Richard S. Mitchell, Bian Tan, Nancy E. Jordan, Ann F. Johnson, Wilson Baker, Cecil R Slaughter, Marc Minno. States and Counties: Florida: Baker, Calhoun, Citrus, Columbia, Duval, Franklin, Gadsden, Gulf, Jackson, Jefferson, Lafayette, Leon, Liberty, Madison, Okaloosa, Wakulla, Walton. Georgia: Grady, Thomas. Compiled by Tall Timbers Research Station and Land Conservancy.
 
 
Radford, Albert E., Harry E. Ahles, and C. Ritchie Bell. Manual of the Vascular Flora of the Carolinas. 1964, 1968. The University of North Carolina Press. 897. Print.
 

Revision as of 09:53, 9 August 2016

Trichostema dichotomum
Trichostema dichotomum Gil.jpg
Photo taken by Gil Nelson
Scientific classification
Kingdom: Plantae
Division: Magnoliophyta – Flowering plants
Class: Magnoliopsida – Dicotyledons
Order: Lamiales
Family: Lamiaceae ⁄ Labiatae
Genus: Trichostema
Species: T. dichotomum
Binomial name
Trichostema dichotomum
L.
TRIC DICH dist.jpg.jpg
Natural range of Trichostema dichotomum from USDA NRCS Plants Database.

Common name: forked bluecurls

Taxonomic notes

Synonyms: Trichostema dichotomum var. dichotomum; Trichostema dichotomum var. puberulum Fernald & Griscom

Description

"Pubescent annual or weak perennial herbs from tap roots, freely branched with opposite branches. Stems obscurely angled, usually stipitate-glandular, leafy. Leaves entire or toothed, rarely lobed, petiolate to essentially sessile. Inflorescence a panicle of paired, bracteate, helicoid cymes with a flower at the axis of each pair; bracts similar to the leaves, smaller. Calyx bilabiate, lower lip longer than the upper with 3 essentially equal teeth, upper with 2 similar teeth; corolla blue to violet, zygomorphic, 5-lobed, lowest lobe lip-like, 5-10 mm long, lateral triangular-ovate, 2-4 mm long, tube short. Stamens 4, ca. 2X as long as the corolla, curved between the lateral lobes and curled downward almost meeting the lip; stigma bilobed, style curled in same fashion as stamens. Mericarps brownish to olive or blackish, dull, obovoid." < ref name="Radford et al 1964">Radford, Albert E., Harry E. Ahles, and C. Ritchie Bell. Manual of the Vascular Flora of the Carolinas. 1964, 1968. The University of North Carolina Press. 897. Print.</ref>

"At least the principal stems spreading pubescent, the trichomes 0.2-1 mm long, usually bearded at the nodes. Leaves elliptic to elliptic-lanceolate, 2-7 cm long, 0.4-2.3 cm wide, entire, crenate, or rarely loved, base tapered to cuneate or attenuate; petioles 0-1.5 cm long. Mericarps distinctly reticulate, 1.9-2.3 mm long." [1]

Distribution

Ecology

Habitat

In the Coastal Plain in Florida and Georgia, T. dichotomum has been found in burned upland pinewoods; longleaf pine forests; annually burned pine savannas; loamy sand of woodlands; loamy sand of open pine-hickory woods; floodplain forests; sandy loam along bluffs; amidst cypress knees on the shores of lakes and ponds; sandy ridges; burned pond pine shrub bog flatwoods; longleaf pine-turkey oak sand ridges; and at the edges of an open limestone glade. [2] It has been recorded in disturbed habitats such as roadsides; deciduous flatwoods clearings; and a mowed meadow. Associated species include Vaccinium stamineum, Trichostema setaceum, Scoparia, Panicum, Andropogon gerardii, Angelica dentata, and Helianthus radula. [2]

Phenology

It has been recorded flowering September and October, and fruiting September through December. [2]

Seed dispersal

According to Kay Kirkman, a plant ecologist, this species disperses by gravity. [3]

Seed bank and germination

Habitats associated with high light levels and large diurnal temperature fluctuations near tortoise mounds are probably favorable for the establishment of T. dichotomum. [4]

Pollination

The following Hymenoptera families and species were observed visiting flowers of Trichostema dichotomum at Archbold Biological Station: [5]

Halictidae: Lasioglossum placidensis

Use by animals

It was found to be one of the most common spring recruits near recently abandoned tortoise mounds. [6] Bee species, Caupolicana electa and Dialictus placidensis, have been observed on T. dichotomum. [7]

Conservation and management

Cultivation and restoration

Photo Gallery

References and notes

  1. Cite error: Invalid <ref> tag; no text was provided for refs named Radford et al 1964
  2. 2.0 2.1 2.2 Florida State University Robert K. Godfrey Herbarium database. URL: http://herbarium.bio.fsu.edu. Last accessed: July 2015. Collectors: Loran C. Anderson, Robert K. Godfrey, R. Komarek, R. A. Norris, Andre F. Clewell, K. Craddock Burks, David Dixon, O. Lakela, R. Kral, James P. Gillespie, R. L. Lazor, A. H. Curtiss, Roy Jervis, John B. Nelson, R. E. Perdue, Jr., Richard S. Mitchell, Bian Tan, Nancy E. Jordan, Ann F. Johnson, Wilson Baker, Cecil R Slaughter, Marc Minno. States and Counties: Florida: Baker, Calhoun, Citrus, Columbia, Duval, Franklin, Gadsden, Gulf, Jackson, Jefferson, Lafayette, Leon, Liberty, Madison, Okaloosa, Wakulla, Walton. Georgia: Grady, Thomas. Compiled by Tall Timbers Research Station and Land Conservancy.
  3. Kay Kirkman, unpublished data, 2015.
  4. Bazzaz, F. A. 1979. The physiological ecology of plant succession. Annu. Rev. Ecol. Syst., 10:351- 371.
  5. Deyrup, M.A. and N.D. 2015. Database of observations of Hymenoptera visitations to flowers of plants on Archbold Biological Station, Florida, USA.
  6. Kaczor, S. A. and D. C. Hartnett (1990). "Gopher tortoise (gopherus polyphemus) effects on soils and vegetation in a Florida sandhill." American Midland Naturalist 123: 100-111.
  7. Deyrup, M. and L. Deyrup (2012). "The diversity of insects visiting flowers of saw palmetto (Arecaceae)." Florida Entomologist 95(3): 711-730.