Difference between revisions of "Lycopodiella alopecuroides"

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Common name: Foxtail clubmoss
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Common name: Foxtail clubmoss<ref name="weakley">Weakley, A.S. 2015. Flora of the southern and mid-atlantic states. Working Draft of 21 May 2015. University of North Carolina at Chapel Hill, Chapel Hill, North Carolina.</ref>
 
==Taxonomic notes==
 
==Taxonomic notes==
Synonym: ''Lycopodium alopecuroides'' L.
+
Synonyms: ''Lycopodium alopecuroides'' Linnaeus.<ref name="weakley">Weakley, A.S. 2015. Flora of the southern and mid-atlantic states. Working Draft of 21 May 2015. University of North Carolina at Chapel Hill, Chapel Hill, North Carolina.</ref>
 +
 
 +
Varieties: none.<ref name="weakley">Weakley, A.S. 2015. Flora of the southern and mid-atlantic states. Working Draft of 21 May 2015. University of North Carolina at Chapel Hill, Chapel Hill, North Carolina.</ref>
 
==Description==  
 
==Description==  
 
<!-- Basic life history facts such as annual/perrenial, monoecious/dioecious, root morphology, seed type, etc. -->
 
<!-- Basic life history facts such as annual/perrenial, monoecious/dioecious, root morphology, seed type, etc. -->
 
A description of ''Lycopodiella alopecuroides'' is provided in [http://efloras.org/florataxon.aspx?flora_id=1&taxon_id=233500755 The Flora of North America].
 
A description of ''Lycopodiella alopecuroides'' is provided in [http://efloras.org/florataxon.aspx?flora_id=1&taxon_id=233500755 The Flora of North America].
  
"Stems subterranean or on the surface, branching dichotomously and ascending or dividing into horizontal and erect branches. Roots adventitious along the creeping stem. Leaves numerous, small, scale-or awn-like, to 13 mm long, appressed or spreading, sometimes partly fused with the branch, simple,  entire or toothed, with a midrib only. Sporophylls more or less like the sterile leaves, either in cauline bands or clustered in sessile or pedunculate, terminal strobili. Sporangia solitary at the leaf base, subglobose to reniform, opening by a transverse slit; spores uniform, tetrahedral. Gametophytes epigeous, green and thalloid, or fleshy, subterranean and mycorrhizal." <ref name="Radford et al 1964">Radford, Albert E., Harry E. Ahles, and C. Ritchie Bell. Manual of the Vascular Flora of the Carolinas. 1964, 1968. The University of North Carolina Press. 3-6. Print.</ref>
+
"Stems subterranean or on the surface, branching dichotomously and ascending or dividing into horizontal and erect branches. Roots adventitious along the creeping stem. Leaves numerous, small, scale-or awn-like, to 13 mm long, appressed or spreading, sometimes partly fused with the branch, simple,  entire or toothed, with a midrib only. Sporophylls more or less like the sterile leaves, either in cauline bands or clustered in sessile or pedunculate, terminal strobili. Sporangia are solitary at the leaf base, subglobose to reniform, opening by a transverse slit; spores uniform, tetrahedral. Gametophytes epigeous, green and thalloid, or fleshy, subterranean and mycorrhizal."<ref name="Radford et al 1964">Radford, Albert E., Harry E. Ahles, and C. Ritchie Bell. Manual of the Vascular Flora of the Carolinas. 1964, 1968. The University of North Carolina Press. 3-6. Print.</ref>
  
"Sterile stems not dorsiventral, arching and rooting at the tip or procumbent and rooting throughout, 2-5 dm long, 3-4 mm in diam.; leaves deciduous except for the evergreen apex (“bud”). Leaves linear-subulate, upcurving, strongly ciliate-denticulate. Fertile stems erect, widely spaced, rarely forking or fascinated, 10-45 cm long, 2-3 mm in diam., 5-13 mm broad including the leaves which are as those of the procumbent branches but longer, ascending, more or less flat when dry, and densely overlapping. Strobili usually solitary, 2-10 cm long, 11-20 mm broad, conspicuously broader than the stem, Sporophylls subulate, spreading, strong ciliate-denticulate. Sporangia globose. Plants without gemmae." <ref name="Radford et al 1964"/>
+
"Sterile stems not dorsiventral, arching and rooting at the tip or procumbent and rooting throughout, 2-5 dm long, 3-4 mm in diam.; leaves deciduous except for the evergreen apex (“bud”). Leaves linear-subulate, upcurving, strongly ciliate-denticulate. Fertile stems erect, widely spaced, rarely forking or fascinated, 10-45 cm long, 2-3 mm in diam., 5-13 mm broad including the leaves which are as those of the procumbent branches but longer, ascending, more or less flat when dry, and densely overlapping. Strobili usually solitary, 2-10 cm long, 11-20 mm broad, conspicuously broader than the stem, Sporophylls subulate, spreading, strong ciliate-denticulate. Sporangia are globose. Plants without gemmae."<ref name="Radford et al 1964"/>
  
 
==Distribution==
 
==Distribution==
 +
This plant ranges from southeast Massachusetts to Florida, and west to eastern Texas. There are disjunct populations in the Cumberland Plateau of Kentucky, Tennessee, and Virginia, the Allegheny Mountains of West Virginia, the eastern Highland Rim of Tennessee, and Maine. There are also populations in southern Mexico through Central America to South America and Cuba.<ref name="weakley">Weakley, A.S. 2015. Flora of the southern and mid-atlantic states. Working Draft of 21 May 2015. University of North Carolina at Chapel Hill, Chapel Hill, North Carolina.</ref>
 
==Ecology==
 
==Ecology==
 
===Habitat=== <!--Natural communities, human disturbed habitats, topography, hydrology, soils, light, fire regime requirements for removal of competition, etc.-->
 
===Habitat=== <!--Natural communities, human disturbed habitats, topography, hydrology, soils, light, fire regime requirements for removal of competition, etc.-->
''L. alopecuroides'' occurs in moist to wet semi-shaded soils. <ref name="FSU Herbarium">Florida State University Robert K. Godfrey Herbarium database. URL: [http://herbarium.bio.fsu.edu http://herbarium.bio.fsu.edu]. Last accessed: June 2014.  Collectors:  F. Almeda, Loran C. Anderson, K. Craddock Burks, A. H. Curtiss, Mary Davis, J. P. Gillespie Robert K. Godfrey, Richard D. Houk, C. Jackson, Michael R. Jenkins, Roy Komarek, R. Kral, O. Lakela, R. Lazor, S. W. Leonard, Sidney McDaniel, John T. Mickel, Leon Neel, James H. Peck, Alan R. Smith, E. Laurence Thurston, L. B. Trott, E.S. Ford, P.L. Redfearn, Jr., Allen G. Shuey, Travis MacClendon, Karen MacClendon, and Floyd Griffith.  States and Counties:  Florida: Alachua, Baker, Bay, Calhoun, Clay, Duval, Franklin, Gulf, Hillsborough, Jackson, Jefferson, Leon, Liberty, Manatee, Okaloosa, Santa Rosa, Wakulla, and Washington.  Georgia: Grady and Thomas.</ref> Typical soil types range from sandy-peaty soils to loamy sand. <ref name="FSU Herbarium"/> This species can be found in several native habitats, but also in disturbed areas. Native habitat types include wet pine savanna, <ref name="Brewer et al 2011">Brewer, J. S., D. J. Baker, et al. (2011). "Carnivory in plants as a beneficial trait in wetlands." Aquatic Botany 94: 62-70.</ref> hillside bogs, seasonally inundated meadows, damp grassy palmetto woods, and on the edges of ponds, creeks and titi thickets. <ref name="FSU Herbarium"/> It also occurs in disturbed habitat like powerline corridors, roadsides, railways, pine plantations, and borrow pits. <ref name="FSU Herbarium"/> Associated species include ''Ilex myrtifolia, Nyssa biflora, Pinus palutris, Aristida stricta, Cyrilla racemiflora, Juncus, Sarracenia, Rhynchospora, Xyris, Eriocaulon, Liquidambar styraciflua, Quercus laevis, Pinguicula, Kalmia hirsuta, Serenoa repens, Hypericum, Cliftonia,'' and ''Magnolia virginiana''. <ref name="FSU Herbarium"/>
+
''L. alopecuroides'' occurs in moist to wet semi-shaded soils.<ref name="FSU Herbarium">Florida State University Robert K. Godfrey Herbarium database. URL: [http://herbarium.bio.fsu.edu http://herbarium.bio.fsu.edu]. Last accessed: June 2014.  Collectors:  F. Almeda, Loran C. Anderson, K. Craddock Burks, A. H. Curtiss, Mary Davis, J. P. Gillespie Robert K. Godfrey, Richard D. Houk, C. Jackson, Michael R. Jenkins, Roy Komarek, R. Kral, O. Lakela, R. Lazor, S. W. Leonard, Sidney McDaniel, John T. Mickel, Leon Neel, James H. Peck, Alan R. Smith, E. Laurence Thurston, L. B. Trott, E.S. Ford, P.L. Redfearn, Jr., Allen G. Shuey, Travis MacClendon, Karen MacClendon, and Floyd Griffith.  States and Counties:  Florida: Alachua, Baker, Bay, Calhoun, Clay, Duval, Franklin, Gulf, Hillsborough, Jackson, Jefferson, Leon, Liberty, Manatee, Okaloosa, Santa Rosa, Wakulla, and Washington.  Georgia: Grady and Thomas.</ref> Typical soil types range from sandy-peaty soils to loamy sand.<ref name="FSU Herbarium"/> This species can be found in several native habitats, but also in disturbed areas. Native habitat types include wet pine savanna,<ref name="Brewer et al 2011">Brewer, J. S., D. J. Baker, et al. (2011). "Carnivory in plants as a beneficial trait in wetlands." Aquatic Botany 94: 62-70.</ref> hillside bogs, seasonally inundated meadows, damp grassy palmetto woods, and on the edges of ponds, creeks, and titi thickets.<ref name="FSU Herbarium"/> It also occurs in disturbed habitat like powerline corridors, roadsides, railways, pine plantations, and borrow pits.<ref name="FSU Herbarium"/> Associated species include ''Ilex myrtifolia, Nyssa biflora, Pinus palutris, Aristida stricta, Cyrilla racemiflora, Juncus, Sarracenia, Rhynchospora, Xyris, Eriocaulon, Liquidambar styraciflua, Quercus laevis, Pinguicula, Kalmia hirsuta, Serenoa repens, Hypericum, Cliftonia,'' and ''Magnolia virginiana''.<ref name="FSU Herbarium"/>
  
 
<!--===Phenology===--> <!--Timing off flowering, fruiting, seed dispersal, and environmental triggers.  Cite PanFlora website if appropriate: http://www.gilnelson.com/PanFlora/ -->
 
<!--===Phenology===--> <!--Timing off flowering, fruiting, seed dispersal, and environmental triggers.  Cite PanFlora website if appropriate: http://www.gilnelson.com/PanFlora/ -->
Line 38: Line 41:
 
<!--===Seed bank and germination===-->
 
<!--===Seed bank and germination===-->
 
===Fire ecology=== <!--Fire tolerance, fire dependence, adaptive fire responses-->
 
===Fire ecology=== <!--Fire tolerance, fire dependence, adaptive fire responses-->
This species occurs in habitat that is maintained by frequent or annual fire. <ref name="FSU Herbarium"/>
+
This species occurs in habitat that is maintained by frequent or annual fire.<ref name="FSU Herbarium"/>
  
 
===Pollination===  
 
===Pollination===  
The following Hymenoptera families and species were observed visiting flowers of ''Lycopodiella alopecuroides'' at Archbold Biological Station: <ref name="Deyrup 2015">Deyrup, M.A. and N.D. 2015. Database of observations of Hymenoptera visitations to flowers of plants on Archbold Biological Station, Florida, USA.</ref>
+
The following Hymenoptera families and species were observed visiting flowers of ''Lycopodiella alopecuroides'' at Archbold Biological Station:<ref name="Deyrup 2015">Deyrup, M.A. and N.D. 2015. Database of observations of Hymenoptera visitations to flowers of plants on Archbold Biological Station, Florida, USA.</ref>
  
 
Halictidae:  ''Augochlorella aurata''
 
Halictidae:  ''Augochlorella aurata''

Revision as of 15:55, 25 September 2020

Lycopodiella alopecuroides
Lycopodiella alopecuroides Gil.jpg
Photo taken by Gil Nelson
Scientific classification
Kingdom: Plantae
Division: Lycopodiophyta – Lycopods
Class: Lycopodiopsida
Order: Lycopodiales
Family: Lycopodiaceae
Genus: Lycopodiella
Species: L. alopecuroides
Binomial name
Lycopodiella alopecuroides
(L.) Cranfill
LYCO ALOP dist.jpg
Natural range of Lycopodiella alopecuroides from USDA NRCS Plants Database.

Common name: Foxtail clubmoss[1]

Taxonomic notes

Synonyms: Lycopodium alopecuroides Linnaeus.[1]

Varieties: none.[1]

Description

A description of Lycopodiella alopecuroides is provided in The Flora of North America.

"Stems subterranean or on the surface, branching dichotomously and ascending or dividing into horizontal and erect branches. Roots adventitious along the creeping stem. Leaves numerous, small, scale-or awn-like, to 13 mm long, appressed or spreading, sometimes partly fused with the branch, simple, entire or toothed, with a midrib only. Sporophylls more or less like the sterile leaves, either in cauline bands or clustered in sessile or pedunculate, terminal strobili. Sporangia are solitary at the leaf base, subglobose to reniform, opening by a transverse slit; spores uniform, tetrahedral. Gametophytes epigeous, green and thalloid, or fleshy, subterranean and mycorrhizal."[2]

"Sterile stems not dorsiventral, arching and rooting at the tip or procumbent and rooting throughout, 2-5 dm long, 3-4 mm in diam.; leaves deciduous except for the evergreen apex (“bud”). Leaves linear-subulate, upcurving, strongly ciliate-denticulate. Fertile stems erect, widely spaced, rarely forking or fascinated, 10-45 cm long, 2-3 mm in diam., 5-13 mm broad including the leaves which are as those of the procumbent branches but longer, ascending, more or less flat when dry, and densely overlapping. Strobili usually solitary, 2-10 cm long, 11-20 mm broad, conspicuously broader than the stem, Sporophylls subulate, spreading, strong ciliate-denticulate. Sporangia are globose. Plants without gemmae."[2]

Distribution

This plant ranges from southeast Massachusetts to Florida, and west to eastern Texas. There are disjunct populations in the Cumberland Plateau of Kentucky, Tennessee, and Virginia, the Allegheny Mountains of West Virginia, the eastern Highland Rim of Tennessee, and Maine. There are also populations in southern Mexico through Central America to South America and Cuba.[1]

Ecology

Habitat

L. alopecuroides occurs in moist to wet semi-shaded soils.[3] Typical soil types range from sandy-peaty soils to loamy sand.[3] This species can be found in several native habitats, but also in disturbed areas. Native habitat types include wet pine savanna,[4] hillside bogs, seasonally inundated meadows, damp grassy palmetto woods, and on the edges of ponds, creeks, and titi thickets.[3] It also occurs in disturbed habitat like powerline corridors, roadsides, railways, pine plantations, and borrow pits.[3] Associated species include Ilex myrtifolia, Nyssa biflora, Pinus palutris, Aristida stricta, Cyrilla racemiflora, Juncus, Sarracenia, Rhynchospora, Xyris, Eriocaulon, Liquidambar styraciflua, Quercus laevis, Pinguicula, Kalmia hirsuta, Serenoa repens, Hypericum, Cliftonia, and Magnolia virginiana.[3]

Fire ecology

This species occurs in habitat that is maintained by frequent or annual fire.[3]

Pollination

The following Hymenoptera families and species were observed visiting flowers of Lycopodiella alopecuroides at Archbold Biological Station:[5]

Halictidae: Augochlorella aurata

Conservation and management

Cultivation and restoration

Photo Gallery

References and notes

  1. 1.0 1.1 1.2 1.3 Weakley, A.S. 2015. Flora of the southern and mid-atlantic states. Working Draft of 21 May 2015. University of North Carolina at Chapel Hill, Chapel Hill, North Carolina.
  2. 2.0 2.1 Radford, Albert E., Harry E. Ahles, and C. Ritchie Bell. Manual of the Vascular Flora of the Carolinas. 1964, 1968. The University of North Carolina Press. 3-6. Print.
  3. 3.0 3.1 3.2 3.3 3.4 3.5 Florida State University Robert K. Godfrey Herbarium database. URL: http://herbarium.bio.fsu.edu. Last accessed: June 2014. Collectors: F. Almeda, Loran C. Anderson, K. Craddock Burks, A. H. Curtiss, Mary Davis, J. P. Gillespie Robert K. Godfrey, Richard D. Houk, C. Jackson, Michael R. Jenkins, Roy Komarek, R. Kral, O. Lakela, R. Lazor, S. W. Leonard, Sidney McDaniel, John T. Mickel, Leon Neel, James H. Peck, Alan R. Smith, E. Laurence Thurston, L. B. Trott, E.S. Ford, P.L. Redfearn, Jr., Allen G. Shuey, Travis MacClendon, Karen MacClendon, and Floyd Griffith. States and Counties: Florida: Alachua, Baker, Bay, Calhoun, Clay, Duval, Franklin, Gulf, Hillsborough, Jackson, Jefferson, Leon, Liberty, Manatee, Okaloosa, Santa Rosa, Wakulla, and Washington. Georgia: Grady and Thomas.
  4. Brewer, J. S., D. J. Baker, et al. (2011). "Carnivory in plants as a beneficial trait in wetlands." Aquatic Botany 94: 62-70.
  5. Deyrup, M.A. and N.D. 2015. Database of observations of Hymenoptera visitations to flowers of plants on Archbold Biological Station, Florida, USA.