Difference between revisions of "Viola palmata"
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==Ecology== | ==Ecology== | ||
===Habitat=== <!--Natural communities, human disturbed habitats, topography, hydrology, soils, light, fire regime requirements for removal of competition, etc.--> | ===Habitat=== <!--Natural communities, human disturbed habitats, topography, hydrology, soils, light, fire regime requirements for removal of competition, etc.--> | ||
| − | In the Coastal Plain in Florida and Georgia, ''V. palmata'' has been found in natural longleaf stands; longleaf pine-wiregrass savannas; loamy soil along mesic hardwood slopes; seepages in ravines; river banks; sandy loam under beech-magnolia forests; hardwood hammocks; oak-palmetto woods; alluvial soil of floodplain forests; between road and slash pine/wiregrass flatwoods; recently burned longleaf pine; longleaf pine-palmetto flats; loamy sand of grassy savanna; second growth loblolly pinewoods; sandy soil of turkey oak-longleaf pine flats; calcareous bluffs; upland mixed woodlands; beech-magnolia forests; and loamy humus of shallow depression of shaded limestone rock in hardwood maritime hammock.<ref name="FSU Herbarium">Florida State University Robert K. Godfrey Herbarium database. URL: [http://herbarium.bio.fsu.edu http://herbarium.bio.fsu.edu]. Last accessed: July 2015. Collectors: Loran C. Anderson, K. Craddock Burks, Robert K. Godfrey, S. W. Leonard, Wilson Baker, James R. Burkhalter, Robert Kral, L. B. Trott, J. P. Gillespie, D. B. Ward, J. Beckner, Patricia Elliot, Cecil R Slaughter, S. R. Harrison, C. Jackson, Nir L. Gil, Angus Gholson, Steve L. Orzell, Edwin L. Bridges, A. G. Shuey, Sidney McDaniel, S. C. Hood, Richard Carter, Mark A Garland, Richard S. Mitchell, Karen MacClendon, B. & M. Boothe, M. R. Darst, Rodie White, R. A. Norris, R. Komarek. States and Counties: Florida: Calhoun, Clay, Flagler, Franklin, Gadsden, Hamilton, Holmes, Jackson, Jefferson, Leon, Liberty, Madison, Manatee, Marion, Okaloosa, Santa Rosa, Seminole, St. Johns, Taylor, Wakulla, Washington. Georgia: Thomas. Compiled by Tall Timbers Research Station and Land Conservancy.</ref> <ref name="K04"> Kirkman, L. K., K. L. Coffey, et al. (2004). "Ground cover recovery patterns and life-history traits: implications for restoration obstacles and opportunities in a species-rich savanna." Journal of Ecology 92: 409-421. </ref> It has been found to be extremely vulnerable to disturbance <ref name="K04"/> however has been found growing along roadsides, sands of disturbed hillside bogs, lakesides, calcareous banks between earth road and drainage canal, ectone between oak woodland and open powerline corridor, heavily cattle grazed palmetto flatwoods, and a powerline transect through a longleaf pine-wiregrass savanna. <ref name="FSU Herbarium"/> When exposed to soil disturbance by military training in West Georgia, ''V. palmata'' responds negatively by way of absence.<ref>Dale, V.H., S.C. Beyeler, and B. Jackson. (2002). Understory vegetation indicators of anthropogenic disturbance in longleaf pine forests at Fort Benning, Georgia, USA. Ecological Indicators 1(3):155-170.</ref> | + | In the Coastal Plain in Florida and Georgia, ''V. palmata'' has been found in natural longleaf stands; longleaf pine-wiregrass savannas; loamy soil along mesic hardwood slopes; seepages in ravines; river banks; sandy loam under beech-magnolia forests; hardwood hammocks; oak-palmetto woods; alluvial soil of floodplain forests; between road and slash pine/wiregrass flatwoods; recently burned longleaf pine; longleaf pine-palmetto flats; loamy sand of grassy savanna; second growth loblolly pinewoods; sandy soil of turkey oak-longleaf pine flats; calcareous bluffs; upland mixed woodlands; beech-magnolia forests; and loamy humus of shallow depression of shaded limestone rock in hardwood maritime hammock.<ref name="FSU Herbarium">Florida State University Robert K. Godfrey Herbarium database. URL: [http://herbarium.bio.fsu.edu http://herbarium.bio.fsu.edu]. Last accessed: July 2015. Collectors: Loran C. Anderson, K. Craddock Burks, Robert K. Godfrey, S. W. Leonard, Wilson Baker, James R. Burkhalter, Robert Kral, L. B. Trott, J. P. Gillespie, D. B. Ward, J. Beckner, Patricia Elliot, Cecil R Slaughter, S. R. Harrison, C. Jackson, Nir L. Gil, Angus Gholson, Steve L. Orzell, Edwin L. Bridges, A. G. Shuey, Sidney McDaniel, S. C. Hood, Richard Carter, Mark A Garland, Richard S. Mitchell, Karen MacClendon, B. & M. Boothe, M. R. Darst, Rodie White, R. A. Norris, R. Komarek. States and Counties: Florida: Calhoun, Clay, Flagler, Franklin, Gadsden, Hamilton, Holmes, Jackson, Jefferson, Leon, Liberty, Madison, Manatee, Marion, Okaloosa, Santa Rosa, Seminole, St. Johns, Taylor, Wakulla, Washington. Georgia: Thomas. Compiled by Tall Timbers Research Station and Land Conservancy.</ref> <ref name="K04"> Kirkman, L. K., K. L. Coffey, et al. (2004). "Ground cover recovery patterns and life-history traits: implications for restoration obstacles and opportunities in a species-rich savanna." Journal of Ecology 92: 409-421. </ref> It has been found to be extremely vulnerable to disturbance <ref name="K04"/> however has been found growing along roadsides, sands of disturbed hillside bogs, lakesides, calcareous banks between earth road and drainage canal, ectone between oak woodland and open powerline corridor, heavily cattle grazed palmetto flatwoods, and a powerline transect through a longleaf pine-wiregrass savanna. <ref name="FSU Herbarium"/> When exposed to soil disturbance by military training in West Georgia, ''V. palmata'' responds negatively by way of absence.<ref>Dale, V.H., S.C. Beyeler, and B. Jackson. (2002). Understory vegetation indicators of anthropogenic disturbance in longleaf pine forests at Fort Benning, Georgia, USA. Ecological Indicators 1(3):155-170.</ref> It also responds negatively to soil disturbance by agriculture in Southwest Georgia.<ref>Kirkman, L.K., K.L. Coffey, R.J. Mitchell, and E.B. Moser. Ground Cover Recovery Patterns and Life-History Traits: Implications for Restoration Obstacles and Opportunities in a Species-Rich Savanna. (2004). Journal of Ecology 92(3):409-421.</ref> |
Substrate types include loamy soil, loamy sand, sandy loam, peaty sand, alluvial soil, sandy-peaty soil, sandy-clayey soil, limestone and loamy humus. <ref name="FSU Herbarium"/> Associated species include ''Carex crebriflora, Viola floridana, Aristida, Eleocharis, V. villosa, V. lanceolata, Chaptalia, Lycopodium, Sisyrinchium, Calopogon'', and ''Hypericum.'' <ref name="FSU Herbarium"/> | Substrate types include loamy soil, loamy sand, sandy loam, peaty sand, alluvial soil, sandy-peaty soil, sandy-clayey soil, limestone and loamy humus. <ref name="FSU Herbarium"/> Associated species include ''Carex crebriflora, Viola floridana, Aristida, Eleocharis, V. villosa, V. lanceolata, Chaptalia, Lycopodium, Sisyrinchium, Calopogon'', and ''Hypericum.'' <ref name="FSU Herbarium"/> | ||
Revision as of 14:50, 15 July 2019
| Viola palmata | |
|---|---|
| |
| Photo was taken by Gil Nelson | |
| Scientific classification | |
| Kingdom: | Plantae |
| Division: | Magnoliophyta – Flowering plants |
| Class: | Magnoliopsida – Dicotyledons |
| Order: | Violales |
| Family: | Violaceae |
| Genus: | Viola |
| Species: | V. palmata |
| Binomial name | |
| Viola palmata L. (pro sp.) | |
| |
| Natural range of Viola palmata from USDA NRCS Plants Database. | |
Common names: Early blue violet, Wood violet
Contents
Taxonomic notes
Synonyms: Viola palmata var. palmata; Viola palmata var. triloba (Schweinitz) Gingins ex A.P. de Candolle; Viola triloba Schweinitz var. triloba; Viola chalcosperma Brainerd; Viola ×palmata Linnaeus (pro sp.); Viola triloba Schweinitz; V. stoneana
Description
"Herbaceous, rhizomatous or stoloniferous perennials, or winter annuals. Leaves crenate or crenate-serrate, dissected, lobed or unlobed; petiolate; stipules conspicuous. Flowers zygomorphic, peduncles directly from the rhizome (acaulescent) or peduncles axillary (caulescent). Chasmogamous flowers with lateral petals often bearded, lower petal spurred; anthers usually seated in the throat, orange appendages conspicuous, lower stamens spurred; spurs fitting in to the spur of the corolla; styles usually clavate, variously shaped at the apex. Dates for Viola, except V. pedata, are for Chasmogamous flowering only, cleistogamous flowering and fruiting commence shortly after chasmogamous flowering and continues until frost. Hybrids are so numerous in this genus that space is not taken to list them." [1]
"Plant acaulescent; rhizomes elongate, stocky, usually more than 4 mm in diam., horizontal, often freely branched. Leaves widely ovate, 5-10 cm long, or occasionally larger, crenate, unlobed, lobed or dissected, cordate, glabrous or pubescent; petioles to 2 dm long; stipules linear to linear-lanceolate, 0.5-2 cm long, fimbriate, rarely entire. Peduncles of chasmogamous flowers nearly as long as the leaves, or slightly longer. Chasmogamous flowers 1.5-4 cm broad, not flat, petals deep reddish violet, lateral petals bearded with clavate trichomes, spurred petal usually glabrous, or occasionally with some trichomes; sepals 6-10 mm long, acute to obtuse, glabrous to ciliate or only the auricles ciliate, auricles 0.5-1 mm long; stamen appendages brilliant orange, deep in the throat of the flower; style clavate. Cleistogamous flowers on divergent peduncles 3-12 cm long or the peduncles erect and short; sepals 4-6 mm long, acute, auricles ca. 1 mm long. Capsules 6-12 mm long. Seeds brown, usually mottled, 1.5-2.3 mm long." [1]
Distribution
Ecology
Habitat
In the Coastal Plain in Florida and Georgia, V. palmata has been found in natural longleaf stands; longleaf pine-wiregrass savannas; loamy soil along mesic hardwood slopes; seepages in ravines; river banks; sandy loam under beech-magnolia forests; hardwood hammocks; oak-palmetto woods; alluvial soil of floodplain forests; between road and slash pine/wiregrass flatwoods; recently burned longleaf pine; longleaf pine-palmetto flats; loamy sand of grassy savanna; second growth loblolly pinewoods; sandy soil of turkey oak-longleaf pine flats; calcareous bluffs; upland mixed woodlands; beech-magnolia forests; and loamy humus of shallow depression of shaded limestone rock in hardwood maritime hammock.[2] [3] It has been found to be extremely vulnerable to disturbance [3] however has been found growing along roadsides, sands of disturbed hillside bogs, lakesides, calcareous banks between earth road and drainage canal, ectone between oak woodland and open powerline corridor, heavily cattle grazed palmetto flatwoods, and a powerline transect through a longleaf pine-wiregrass savanna. [2] When exposed to soil disturbance by military training in West Georgia, V. palmata responds negatively by way of absence.[4] It also responds negatively to soil disturbance by agriculture in Southwest Georgia.[5]
Substrate types include loamy soil, loamy sand, sandy loam, peaty sand, alluvial soil, sandy-peaty soil, sandy-clayey soil, limestone and loamy humus. [2] Associated species include Carex crebriflora, Viola floridana, Aristida, Eleocharis, V. villosa, V. lanceolata, Chaptalia, Lycopodium, Sisyrinchium, Calopogon, and Hypericum. [2]
Phenology
V. palmata is a summer forb. [3] It has been recorded flowering January through May, August, September, and in December with peak inflorescence in February and March.[6][2] It has been observed fruiting February, March, April, August and September.[2]
Seed dispersal
V. palmata is thought to be dispersed by ants and/or explosive dehiscence. [7]
Fire ecology
It has been observed flowering February and March after a fire. [2]
Conservation and management
Cultivation and restoration
Photo Gallery
Viola palmata Photo by Kevin Robertson
Thomas County, GA
References and notes
- ↑ 1.0 1.1 Radford, Albert E., Harry E. Ahles, and C. Ritchie Bell. Manual of the Vascular Flora of the Carolinas. 1964, 1968. The University of North Carolina Press. 665. Print.
- ↑ 2.0 2.1 2.2 2.3 2.4 2.5 2.6 Florida State University Robert K. Godfrey Herbarium database. URL: http://herbarium.bio.fsu.edu. Last accessed: July 2015. Collectors: Loran C. Anderson, K. Craddock Burks, Robert K. Godfrey, S. W. Leonard, Wilson Baker, James R. Burkhalter, Robert Kral, L. B. Trott, J. P. Gillespie, D. B. Ward, J. Beckner, Patricia Elliot, Cecil R Slaughter, S. R. Harrison, C. Jackson, Nir L. Gil, Angus Gholson, Steve L. Orzell, Edwin L. Bridges, A. G. Shuey, Sidney McDaniel, S. C. Hood, Richard Carter, Mark A Garland, Richard S. Mitchell, Karen MacClendon, B. & M. Boothe, M. R. Darst, Rodie White, R. A. Norris, R. Komarek. States and Counties: Florida: Calhoun, Clay, Flagler, Franklin, Gadsden, Hamilton, Holmes, Jackson, Jefferson, Leon, Liberty, Madison, Manatee, Marion, Okaloosa, Santa Rosa, Seminole, St. Johns, Taylor, Wakulla, Washington. Georgia: Thomas. Compiled by Tall Timbers Research Station and Land Conservancy.
- ↑ 3.0 3.1 3.2 Kirkman, L. K., K. L. Coffey, et al. (2004). "Ground cover recovery patterns and life-history traits: implications for restoration obstacles and opportunities in a species-rich savanna." Journal of Ecology 92: 409-421.
- ↑ Dale, V.H., S.C. Beyeler, and B. Jackson. (2002). Understory vegetation indicators of anthropogenic disturbance in longleaf pine forests at Fort Benning, Georgia, USA. Ecological Indicators 1(3):155-170.
- ↑ Kirkman, L.K., K.L. Coffey, R.J. Mitchell, and E.B. Moser. Ground Cover Recovery Patterns and Life-History Traits: Implications for Restoration Obstacles and Opportunities in a Species-Rich Savanna. (2004). Journal of Ecology 92(3):409-421.
- ↑ Nelson, G. PanFlora: Plant data for the eastern United States with emphasis on the Southeastern Coastal Plains, Florida, and the Florida Panhandle. www.gilnelson.com/PanFlora/ Accessed: 15 DEC 2016
- ↑ Kirkman, L. Katherine. Unpublished database of seed dispersal mode of plants found in Coastal Plain longleaf pine-grasslands of the Jones Ecological Research Center, Georgia.
